Honestly, I think it's gotta be eramis. Before I explain why, I want to be clear that she is still our enemy bc she is actively working with the witness to achieve the final shape, and her suffering does not relieve her of the responsibility she has for Rasputin's death and the ensuing consequences of the traveler being in orbit and being a target for the witness. However, we can acknowledge this while simultaneously acknowledging the amount of suffering she has endured. Eramis is old. Somehow old enough to remember riis and the eliksni Golden age. She saw her people endure the whirlwind and gained a jaded view on the traveler as she watched her people go from one of technical superiority and utopia to being reduced to shameless pirates scavenging the remains of other dying civilizations. Despite witnessing these bleak events, eramis did not give up. Rather than give into despair, she attempted to do something to make life better for her people. When she caught wind of the darkness and stasis on Europa, she seized this power not to get vengeance on the traveler but to empower her people once again. Can we blame her for this? She wanted to empower her people and rebuild their civilization with this power as the foundation. The vanguard did not want the fallen to have this power in the same way we didn't want Iraq to have wmds in the early 2000s (don't make this political, just a comparison), so they prioritized taking this power away. And rightfully so, eramis had demonstrated her hostility towards us as head of the Scourge of the past eliksni syndicate leading the operation to retrieve black armory tech and also to reclaim siva in zero hour. She had to watch us dismantle her operation and her attempt to rebuild riis on Europa, watching her people fall yet again. We all know how beyond light turned out for her, with her becoming an eliksnicicle for the better part of 2 years. These 2 years were likely the easiest of eramis' life however, as she was spared watching her people subjugated as wrathborn, at least for awhile. When she was freed from her stasis prison, she found herself in a new captivity doing the witness' bidding. In her quest to retrieve the relics of nezarec as well as in the following operation to destroy the traveler through the warsat network, she witnessed the mindless husks of her people that had their personalities hollowed out as wrathborn and converted to expendable tools of the witness. Not only this, but the witness also placed eramis' former mate in her court, resurrected as a scorn. Can you imagine the agony of having to see the love of your life puppeted around in a decaying zombie like state without any semblance of her former personality left? Eramis has watched the eliksni used as tools of greater beings time and time again. We can tell this is beginning to wear on her and cause cognitive dissonance within her as she tried to warn us about the shadow legion trap that ultimately killed Amanda Holliday. She then apparently saved mithrax from the same explosion by blocking the tunnel with stasis crystals and then leaving in peace. Don't get me wrong, eramis is still our enemy and must be eliminated or captured. But it is evident that she at least feels some remorse for what she has done, and also that she has done these things in an attempt to provide for her people. Instead, her people have been subjugated to fates worse than death, and eramis is partially responsible. This guilt has got to cause daily suffering for her greater than just the loss of a dear friend.