Sort of a grey area, but mara sov. The awoken use and were made by both the light and dark. She took out the house of wolves and took on Oryx. Also without her clearing the way we probably couldn't have killed oryx.
Toland is also another contender since he is the only guardian to learn the death singer's song and to navigate the ascendant plan.
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what exactly is Toland right now? Is it something like his disembodied soul floating around? And is he stuck in that form, and bound to the ascendant plane?
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Toldand might have a bit of difficulty showing his destructive power, considering hes an orb
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Just causes hes a disembodied soul doesn't mean he cant do things. He did spawn platforms for us when we first entered the dreaming city. He has also guided us and Eris. Speaking of which Eris is also pretty powerful since she can use hive magic