Theories Hive 4

Any ideas on the lore of 4 phrases Eir, Ur, Xol Yul? I realize each swarm prince representing one and wizards of their spawn. Toland claims to be on a trail of a 5th phrase, "faintly heard from the buzz that once spilled from the shrine" which suggests the Shrine of Oryx, also suggesting the 5th phrase is or relates to Oryx. Toland also suggests that these may be their Gods, or Idols. Or that they may hold key to their ultimate demise, or a bridge to their desires. Thanks guys, please comment any theories or ideas below!